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What is odd.
子句类型: 主语从句 连接词：what。
They have perhaps most benefited from ambition.
子句类型: 表语从句 连接词：that。
if引导状语从句。
What is odd is that they have perhaps most benefited from ambition -- if not always their own then that of their parents and grandparents.
奇怪的是他们也许已经从雄心壮志中获取了最大的利益——如果不是他们自己拥有雄心壮志，那么就是他们父母和祖父母拥有。